Fahmy, M., Andrianoely, D., Wright, P., & Hekkala, E. (2023). Leech-derived iDNA complements traditional surveying methods, enhancing species detections for rapid biodiversity sampling in the tropics.


Hekkala, E. R., Colten, R., Cunningham, S. W. (GSAS), Smith, O., & Ikram, S. (2022). Using Mitogenomes to Explore the Social and Ecological Contexts of Crocodile Mummification in Ancient Egypt. Bulletin of the Peabody Museum of Natural History63(1), 3-14.

Flamio Jr, R. (GSAS), Levano, S. R. (FCRH), Kashiwagi, T., Makkay, A. M. (GSAS), & Hekkala, E. R. (2022). Integrating ecological and molecular data to investigate species maintenance and interspecific hybridization between the redbreast sunfish (Lepomis auritus) and the pumpkinseed sunfish ( gibbosus). Canadian Journal of Zoology, (ja).

Klingler, K., Nichols, L., Hekkala, E., & Peacock, M. (2022). Life on the edge-a changing genetic landscape within an iconic American pika metapopulation over the last half century. Research Square pre-print


Rakotozandry, R., Ranivoharimana, L., Ranivosoa, V., Rasolofomanana, N., Hekkala, E. R. (Fordham), Samonds, K. (2021). Description of the subfossil crocodylians from a new Late Pleistocene subfossil site (Tsaramody, Sambaina Basin, central Madagascar). Malagasy Nature, 15(MN15).

Godfrey, L., Crowley, B., Muldoon, K., Burns, S., Klukkert, Z., Ranivoharimanana, L., Alumbaugh, J., Borths, M., Faina, P., Goodman, S., Guitierrez, I., Hansford, J., Hekkala, E. R. (Fordham), Lewis, M., McGee, D., Perez, V., Rahantaharivao, N., Rasolonjatovo, H., Samonds, K., Scroxton, N., Turvey, S. (2021). Teasing apart impacts of human activity and regional drought on Madagascar’s megafauna: insights from new excavations at Tsimanampesotse and Antsirafaly. Frontiers in Ecology and Evolution.

Williams, K.M., Barkdull, M., Fahmy, M., Hekkala, E., Siddall, M.E. and Kvist, S., 2020. Caught red handed: iDNA points to wild source for CITES-protected contraband leeches. European Journal of Wildlife Research, 66(5), pp.1-10.

Hekkala, E., Gatesy, J., Narechania, A., Meredith, R., Russello, M., Aardema, M.L., Jensen, E., Montanari, S., Brochu, C., Norell, M. and Amato, G., 2021. Paleogenomics illuminates the evolutionary history of the extinct Holocene “horned” crocodile of Madagascar, Voay robustus. Voay_CommsBio2021Communications biology, 4(1), pp.1-11.

Godfrey, L.R., Crowley, B., Muldoon, K.M., Burns, S.J., Scroxton, N., Klukkert, Z.S., Ranivoharimanana, L., Alumbaugh, J., Borths, M. and Vasey, N.,et al. 2021. Teasing Apart Impacts of Human Activity and Regional Drought on M Frontiers in Ecology and Evolution, 9.


ER Hekkala, ML Aardema, A Narechania, G Amato, S Ikram, MH Shirley, …2020. The secrets of Sobek–A crocodile mummy mitogenome from ancient Egypt. Journal of Archaeological Science: Reports 33, 102483.

KM Williams, M Barkdull, M Fahmy, E Hekkala, ME Siddall, S Kvist. 2020. Caught red handed: iDNA points to wild source for CITES-protected contraband leeches. European Journal of Wildlife Research 66 (5), 1-10

Milián-García Y, Amato G, Gatesy J, Hekkala E, Rossi N, Russello M. 2020. Phylogenomics reveals novel relationships among Neotropical crocodiles (Crocodylus spp.). Molecular Phylogenetics and Evolution. 2020 Nov 1;152:106924.


Fahmy, M., Ravelomanantsoa, N.A.F., Youssef, S. Hekkala, E. & Siddall, M. Biological inventory of Ranomafana National Park tetrapods using leech-derived iDNA. Eur J Wildl Res (2019) 65: 70.

Siddall, M. E. , Barkdull, M., Tessler, M., Brugler, M.R., Borda, E., Hekkala, E. (2019) Ideating iDNA: Lessons and limitations from leeches in legacy collections. PLoS ONE 14(2): e0212226.


Milan-Garcia, Y., Russello, M., Castellanos-Labarcena, J., Cichon, M., Kumar, V., Espinos, G., Rose, N., Mazzotti, F., Hekkala, E. R., Amato, G. & Janke, A. (2018). Genetic evidence supports a distinct lineage of American crocodile (Crocodylus acutus) in the Greater Antilles. PeerJ, 6 (E5836).MilanGarcia2018


Peacock, M., M. S., Kirchoff, V., Hekkala, E. R., & Heki, L.G. (2017) Return of a giant:  DNA from archival museum samples helps to identify a unique cutthroat trout lineage formerly thought to be extinct. R. Soc. open sci.4: 171253.

Anco, C., Kolokotronis, S. O., Henschel, P., Cunningham, S. W., Amato, G., & Hekkala, E. R.(2017). Historical mitochondrial diversity in African leopards (Panthera pardus) revealed by archival museum specimens. Mitochondrial DNA Part A, 1-19.Historical mitochondrial diversity in African leopards Panthera pardus revealed by archival museum specimens

Frare, C. F., Matocq, M. D., Feldman, C. R., White, A. M., Manley, P. N., Jermstad, K. D. and Hekkala, E. R. (2017), Landscape disturbance and sporadic hybridization complicate field identification of chipmunks. Jour. Wild. Mgmt.. doi:10.1002/jwmg.21193 frare_et_al-2016-the_journal_of_wildlife_management-2


Peacock, M. M., Gustin, M. S., Kirchoff, V. S., Robinson, M. L., Hekkala, E. R., Pizzarro-Barraza, C., & Loux, T. (2016). Native fishes in the Truckee River: Are in-stream structures and patterns of population genetic structure related? Science of The Total Environment, 563, 221-236.s41559-016-0004peacocketal-2016truckeerivernativefishes

Dowell, S. A., Wood, J. P., Campbell, T. S., Kolokotronis, S. O., & Hekkala, E. R. (2016). Combining genetic and distributional approaches to sourcing introduced species: a case study on the Nile monitor (Varanus niloticus) in Florida. Open Science, 3(4), 150619.dowellsourcingmonitors-full

Dowell, S. A., Portik, D. M., de Buffrénil, V., Ineich, I., Greenbaum, E., Kolokotronis, S. O., & Hekkala, E. R. (2016). Molecular data from contemporary and historical collections reveal a complex story of cryptic diversification in the Varanus (Polydaedalus) niloticus Species Group. Molecular phylogenetics and evolution, 94, 591-604.dowellmpe

Cunningham, S. W., Shirley, M. H., & Hekkala, E. R. (2016). Fine scale patterns of genetic partitioning in the rediscovered African crocodile, Crocodylus suchus (Saint-Hilaire 1807). PeerJ, 4, e1901.cunninghametalpeerj-04-1901

Aplasca, A. C., Iverson, J. B., Welch, M. E., Colosimo, G., & Hekkala, E. R. (2016). Genetic diversity and structure in the Endangered Allen Cays Rock Iguana, Cyclura cychlura inornata. PeerJ, 4, e1793.aplascaetalpeerj-1793

Dowell, S. A., & Hekkala, E. R. (2016). Divergent lineages and conserved niches: using ecological niche modeling to examine the evolutionary patterns of the Nile monitor (Varanus niloticus). Evolutionary Ecology, 30(3), 471-485.dowell-and-hekkala-2016-evol-ecol


Carlson D, R Morse & E Hekkala. 2015. Late Spawning Suckers of New York’s Adirondack Mountains. American Currents, Volume 40(4) 1-14.amercurrentsfall2015

Hekkala ER, SG Platt, JB Thorbjarnarson, TR Rainwater, M Tessler, S Cunningham, C Twomey & G Amato. 2015. Integrating molecular, phenotypic and environmental data to elucidate patterns of crocodile hybridization in Belize. Royal Society Open Science 2 (9), 150409.150619-full

Dowell SA, DM Portik, V De Buffrenil V, I Ineich, E Greenbaum, SO Kolokotronis & E Hekkala. 2015. Molecular data from contemporary and historical collections reveal a complex story of cryptic diversification in the Varanus (Polydaedalus) niloticus Species Group. Molecular Phylogenetics and Evolution. doi:10.1016/j.ympev.2015.10.004 dowellmpe

Dowell SA, V de Buffrénil, SO Kolokotronis & ER Hekkala. 2015. Fine-scale genetic analysis of the exploited Nile monitor (Varanus niloticus) in Sahelian Africa. BMC genetics, 16(1), 32.dowelletalbmc2015


Hekkala, E., M.H. Shirley, G. Amato, J.D. Austin, S. Charter, J. Thorbjarnarson, K.A. Vliet, M.L. Houck, R. DeSalle, and M.J. Blum. (2011) Splitting an ancient icon: Analysis of molecular data uncovers cryptic lineages within widespread populations of the Nile crocodile (Crocodylus niloticus). Molecular Ecology (pdf)

Hekkala, E., R. Saumure, J. Jaeger, H.W. Herrmann,, M. Sredl, D. Bradford, D. Drabeck, & M.J. Blum (2011) Resurrecting an extinct species: Archival DNA, taxonomy, and conservation of the Vegas Valley leopard frog. Conservation Genetics (pdf)

Meredith, R. E. Hekkala, G. Amato, and J. Gatesy (2011) A phylogenetic hypothesis for Crocodylus (Crocodylia) based on mitochondrial DNA: Evidence for a transoceanic voyage from Africa to the New World. Molecular Phylogenetics and Evolution (pdf)


Hekkala, E., G. Amato, R. DeSalle & M. Blum (2010) Molecular assessment of population differentiation and individual assignment potential of Nile crocodile (Crocodylus niloticus) populations. Conservation Genetics (pdf)


Hekkala, E., M. Rakotondratsima, and N. Vasey (2007) Habitat and Distribution of the Ruffed Lemur, Varecia, North of the Bay of Antongil in Northeastern Madagascar. Conservation Genetics (pdf)


Schmitz, A., E. Hekkala, T. Shine, H. Nickel, G. Amato, and W. Böhme (2003) Molecular evidence for species level divergence in West African Nile Crocodiles Crocodylus niloticus (Laurenti, 1786). Comptes rendus – Palevol (link)


Randall, J.A., E. Hekkala, L.D. Cooper, J. Barfield (2002) Familiarity and flexible mating strategies of a solitary rodent, Dipodomys ingens. Animal Behavior (pdf)


Randall, J.A., S.M. Hatch, E. Hekkala (1995) Interspecific variation in anti-predator behavior in sympatric species of kangaroo rat. Behavioral Ecology and Sociobiology (pdf)